古地理学报Issue(6)：861-877,17.DOI:10.7605/gdlxb.2014.06.069
志留纪-早泥盆世维管植物的系统发育、幕式演化和地理分布：植物大化石证据
Phylogeny,episodic evolution and geographic distribution of the Silurian-Early Devonian vascular plants:Evidences from plant megafossils
摘要
Abstract
Much progress has been made in the understanding of the phylogeny of Silurian-Early De-vonian vascular land plants during the past years.However,no consensus has been reached considering the delimitation and subdivision of polysporangiophytes,rhyniopsids,and zosterophyllopsids;the sister-group relationship between zosterophyllopsids and lycopsids is questioned;detailed studies of basal eu-phyllophytes are needed.The evolutionary history of the Silurian-Early Devonian vascular plants can be di-vided into two phases:The pre-Pragian prelude phase and the Early Devonian Pragian explosion (burst) phase.The first phase is characterized by initial emergences of representatives of vascular plant major clades.The second is the main phase which witnessed explosive radiation of vascular plant clades.During the Pragian, typical features of lycopsids such as regular phyllotaxy, sporophylls, and strobilus-like structures appeared;the diversification of euphyllophyte lineages occurred;megaphylls and their homolo-gous structures (branch-leaf complexes) evolved multiple times independently;and developmental inter-relationship between sporangia and leafy structures convergently established in different lineages.In South China of the northeastern Gondwanan phytogeographic unit,zosterophyllopsids were dominated by those with spirally arranged sporangia and euphyllophytes showed high diversity and disparity during the Pragian, while in the Laurussian phytogeographic unit,zosterophyllopsids with rowed sporangia were abundant and the appearance of comparable euphyllophytes occurred later than in South China.The cladogenesis of hor-neophytopsids,cooksonioids,and rhynialeans might have mainly occurred at middle and high paleolati-tudes such as southern Laurussia and northwestern Gondwana.The cladogenesis and biogeographic radia-tion of zosterophyllopsids,lycopsids,and euphyllophytes may accord with the “out of the tropics” mod-el,that is,these groups diversified and rapidly radiated earlier at low paleolatitudes;the explosion of vascular plant clades in Pragian was mainly contributed by the evolutionary novelty of tropical floras.关键词
早期维管植物/华南/分支演化/辐射/多样性Key words
early vascular plant/South China/cladogenesis/radiation/diversity分类
生物科学引用本文复制引用
薛进庄,郝守刚..志留纪-早泥盆世维管植物的系统发育、幕式演化和地理分布：植物大化石证据[J].古地理学报,2014,(6):861-877,17.基金项目
国家自然科学基金项目（41272018）、中国地质调查局项目（1212011120142）联合资助 （）
