We would like to thank the curators and staff of the CHS,KUN,MO,P,PE herbaria that offered their facilities and help in examining herbarium specimens,and thank Dr.Ran Wei(Institute of Botany,Chinese Academy of Sciences)for sample collection,Robbin C.Moran(The New York Botanical Garden)for discussions on the morphology of Dryopteridaceae,improving the manuscript,and providing photographs of several neotropical genera.We also thank the Experimental Center of Molecular Biology of the Germplasm Bank of Wild Species,Kunming Institute of Botany,Chinese Acad-emy of Sciences for suggestion of laboratory work,and the iFlora High Performance Computing Center of the Germplasm Bank of Wild Species(iFlora HPC Center of GBOWS,KIB,CAS)for computing.Madagascar collecting permits were granted to GR by Madagascar National Parks and the"Ministère de l'Environnement,de l'Ecologie et des Forêts".The authors are also grateful for field assistance from CNRE-Madagascar and MBG-Madagascar.The MNHN gives access to the collections in the framework of the RECOLNAT National Research Infrastructure.The study was funded by the National Natural Science Foundation of China(Grant No.31970232)and the National Key Basic Research Program of China(Grant No.2014CB954100). （Institute of Botany,Chinese Academy of Sciences）