This work was supported by the project from the National Natural Science Foundation of China(Grant no.42230411)and the Science&Technology Innovation Project of Laoshan Laboratory(Grant no.LSKJ202203206).We are grateful to Antarctic Great Wall Ecology National Observation and Research Station,Arctic Yellow River Earth System National Observation and Research Station and National Arctic and Antarctic Data Center of Polar Research Institute of China for providing the experimental sites and necessary support.We thank all the teammates during 39th Chinese Antarctic Expedition and 13th Chinese Arctic Expedition,for their great efforts and help in sample collection.We appreciate access to the computing resources provided by IEMB-1,a high-performance computation cluster operated by the Institute of Evolution and Marine Biodiversity,Ocean University of China.We thank the two anonymous reviewers for their comments that further improved this manuscript. （Grant no.42230411）