We thank Yang Qiang,Shao Shuai,Feng Yuhui,Li Yang,Yang Shurui,Yu Peng,and Yang Yuguo(staff of the Paleontological Museum of Liaoning,Shenyang,China)and Yu Dongxiang(student of Shandong University of Science and Technology,Qingdao,China)for their help in field work and fossil preparation.We thank Peng Yu(Zhejiang University,Hangzhou,China)for his help with CT scanning of the tooth.We thank Dr.Ren Xinxin(Institute of Geology,Chinese Academy of Geological Sciences,Beijing,China)for his help in finding literature.We also thank four anonymous reviewers,and editor Xu Xing for their constructive criticisms that greatly improved the quality of this paper.This work was supported by the Scientific Research Foundation of Shenyang Normal University(Grant No.BS202207),Program for Innovative Research Team of Excellent Talents in University of Shandong Province(Grant No.2019KJH004),Taishan Scholar Program of Shandong Province(Grant No.tsqn201812070),Educational Department of Liaoning Province(Grant No.JYTQN2023422),Shandong Provincial Natural Science Foundation(Grant No.ZR2017MD031)and the National Natural Science Foundation of China(Grant Nos.41972025,41688103,42161134003). （staff of the Paleontological Museum of Liaoning,Shenyang,China）